NOVA Season 53 Episode 17 Evolution: On the Run ( Air Date 2026-11-11 ) Genre Documentary category. Overview "Run. Leap. Swim. Pounce. Slither. Fly. Life moves in countless ways – but it wasn’t always this way. When animals first emerged, they clung to the seafloor, until a single twitch sparked a revolution. From the first muscle to the first heartbeat, from lobe fins that pushed life onto land to legs built for speed, movement reshaped the planet.".